1. Department of Cell Biology, Faculty of Medicine, Kyoto University, Sakyo-ku, Kyoto 606-8501, Japan
2. Department of Neurosurgery, Faculty of Medicine, Kyoto University, Sakyo-ku, Kyoto 606-8507, Japan
3. KAN Research Institute, Kyoto Research Park, Shimogyo-ku, Kyoto 606-8317, Japan
4. Department of Physiology, Kyoto Prefectural University of Medicine, Kamigyo-ku, Kyoto 602-0841, Japan
5. Department of Molecular Cell Biology, Institute of DNA Medicine, Jikei University School of Medicine, Minato-ku, Tokyo 105-8461, Japan