1. Department of Anatomy, Ajou University School of Medicine, Suwon 16499, Korea
2. Department of Biomedical Sciences, Graduate School, Ajou University School of Medicine, Suwon 16499, Korea
3. These authors contributed equally to this work.
4. Research Center, Cell&Brain Co., Ltd., Jeonju 54871, Korea
5. Department of Medical Lifescience, The Catholic University of Korea, College of Medicine, Seoul 06591, Korea