1. Laboratory of Mucosal Immunology, Department of Biomedical Sciences, Seoul National University College of Medicine, Seoul 03080, Korea
2. These authors contributed equally to this work.
3. Department of Anatomy and Cell Biology, Seoul National University College of Medicine, Seoul 03080, Korea
4. Institute of Allergy and Clinical Immunology, Seoul National University Medical Research Center, Seoul 03080, Korea
5. Department of Biological Sciences, Sungkyunkwan University, Suwon 16419, Korea
6. Biomedical Institute for Convergence at SKKU, Sungkyunkwan University, Suwon 16419, Korea
7. Department of Pathology, Seoul National University College of Medicine, Seoul 03080, Korea
8. Laboratory of Immune Regulation, Department of Biomedical Sciences, Seoul National University College of Medicine, Seoul 03080, Korea
9. Department of Internal Medicine, Seoul National University College of Medicine, Seoul 03080, Korea