1. a School of Conservation Sciences , Bournemouth University, Talbot Campus , Fern Barrow, Poole, Dorset BH12 5BB, UK
2. b School of Geography , University of Southampton , Highfield, Southampton, Hampshire SO17 1BJ, UK
3. c Centre for Earth and Environmental Research , School of Earth Sciences and Geography , Kingston University , Penrhyn Road, Kingston upon Thames, Surrey KT1 2EE, UK