1. Department of Botany and Microbiology, College of Science, King Saud University, Riyadh, Saudi Arabia
2. National Center for Biotechnology, King Abdulaziz City for Science and Technology, Saudi Arabia
3. Department of plant protection, College of Agriculture, King Saud University, Riyadh, Saudi Arabia
4. Department of Molecular Biology and Biotechnology, University of Sheffield, Sheffield, UK