1. Guizhou Academy of Forestry, Guiyang, Guizhou, China
2. Key Laboratory of National Forestry and Grassland Administration on Biodiversity Conservation in Karst Mountainous Areas of Southwestern China, Guiyang, Guizhou, China
3. Advanced Technology and Materials Research Institute, Guizhou Academy of Sciences, Guiyang, Guizhou, China
4. College of Forestry, Guizhou University, Guiyang, Guizhou, China