1. Key Laboratory of Aquaculture in South China Sea for Aquatic Economic Animal of Guangdong Higher Education Institutes, Fisheries College, Guangdong Ocean University, Zhanjiang, China;
2. Guangdong Provincial Key Laboratory of Pathogenic Biology and Epidemiology for Aquatic Animals, College of Fishery, Guangdong Ocean University, Zhanjiang, China;
3. Guangdong Key Laboratory of Control for Diseases of Aquatic Economic Animals, Zhanjiang, China;
4. Department of fisheries and Aquatic Resources Management, University for Development Studies, Tamale, Ghana