1. Graduate School of Life and Environmental Sciences, University of Tsukuba, Tsukuba, Ibaraki, Japan
2. College of Biological Sciences, University of Tsukuba, Tsukuba, Ibaraki, Japan
3. Faculty of Life and Environmental Sciences, University of Tsukuba, Tsukuba, Ibaraki, Japan
4. PRESTO, Japan Science and Technology Agency, Kawaguchi, Saitama, Japan