1. Science for Life Laboratory, Department of Medical Sciences, Uppsala University, Uppsala, Sweden
2. Science for Life Laboratory, Department of Medical Biochemistry and Microbiology, Uppsala University, Uppsala, Sweden
3. Department of Rheumatology, and Department of Biomedical and Clinical Sciences, Linköping University, Linköping, Sweden
4. Department of Biomedical and Clinical Sciences, Linköping University, Linköping, Sweden
5. Broad Institute of MIT and Harvard, Cambridge, MA, USA