1. State Key Laboratory of Biogeology and Environmental Geology, China University of Geosciences, Beijing, China
2. School of the Earth Sciences and Resources, China University of Geosciences, Beijing, China
3. The University of Queensland, Brisbane, Queensland, Australia
4. Institute of Sedimentary Geology, Chengdu University of Technology, Chengdu, China
5. Dinosaur Trackers Research Group, University of Colorado Denver, Denver, CO, USA