1. Graduate School of Bioagricultural Sciences, Nagoya University, Nagoya, Japan
2. Hendrix Genetics BU Layers, Boxmeer, The Netherlands
3. Graduate School of Integrated Sciences for Life, Hiroshima University, Higashi-Hiroshima, Japan
4. Japanese Avian Bioresource Project Research Center, Hiroshima University, Higashi-Hiroshima, Japan