1. Avian Disease Laboratory, College of Veterinary Medicine, Konkuk University, Seoul, Republic of Korea
2. Wildlife Health Laboratory, College of Veterinary Medicine, Konkuk University, Seoul, Republic of Korea
3. KHAV Co, Ltd, Seoul, Republic of Korea
4. Konkuk University Zoonotic Disease Research Center, Seoul, Republic of Korea
5. Department of Epidemiology and Biostatistics, College of Public Health, Institute of Bioinformatics, Center for the Ecology of Infectious Disease, Center for Applied Pathogen Epidemiology and Outbreak Response, University of Georgia, Athens, GA, USA