1. Department of Biodiversity, Faculty of Science and Agriculture, University of Limpopo, Sovenga, Limpopo, South Africa
2. South African Institute for Aquatic Biodiversity, Grahamstown, South Africa
3. School of Life Sciences, University of KwaZulu-Natal, Durban, South Africa
4. Department of Paraclinical Sciences, Faculty of Veterinary Science, University of Pretoria, Onderstepoort, South Africa