1. Department of Biological Sciences, University of Cape Town, Cape Town, South Africa
2. Centre for Statistics in Ecology, Environment and Conservation, Department of Biological Sciences, University of Cape Town, Cape Town, South Africa
3. South African National Biodiversity Institute (SANBI), Cape Town, South Africa
4. Department of Oceanography, Institute for Coastal and Marine Research, Nelson Mandela University, Gqeberha, South Africa